Published Work

Below is a list of publications to which I have contributed:

Limosa 92.2

Our article about the mixed breeding pair Herring Gull x Lesser Black-backed Gull which was breeding in the IJmuiden Forteiland colony in 2017 was published in Dutch scientific magazine Limosa, volume 92.2, in 2019.

Summary (source:

COTTAAR F, VAN KLEINWEE M & VERBEEK-COTTAAR J (2019) Mixed breeding pair Herring Gull Larus argentatus x Lesser Black-backed Gull Larus fuscus. LIMOSA 92 (2): 95-99.

In May 2017, a male Herring Gull was seen together with a female Lesser Black-backed Gull in the gull colony of IJmuiden, the Netherlands. The male had colour-rings green Y.CBC, the female was later colour-ringed with combination green Y.CHA. At the time of the first observation the pair was showing pair bonding behaviour: the male was feeding the female. During the breeding season the pair built a nest and incubated three eggs of which one egg hatched. Unfortunately, the chick only survived for a week. In the previous breeding season the male Herring Gull had been paired with a female (unringed) Herring Gull. Possible reasons to pair up with a female Lesser Black-backed Gull in 2017 include (1) voluntary switching to a new partner because of a previously unsuccessful breeding attempt, (2) forced switching because a previous partner does not return (deceased), (3) a partner is ‘claimed’ by a third individual, or (4) a shortage of partners of the same species (females of large gulls often breed in a colony different than the colony in which they are born). Although the reason for the peculiar mate choice remains unclear, a shortage of Herring Gulls in the colony could have made the male decide to switch to a Lesser Black-backed Gull. From the viewpoint of the female Lesser Black-backed Gull the decision to pair up with a male Herring Gull is intriguing because of the large number of Lesser Black-backed Gulls available in the colony. Documented observations of mixed pairs of Herring Gull x Lesser Black-backed Gull are rare. This was only the second case a mixed breeding pair was discovered in our well-studied gull colony.



Publication: “PAUKŠČIAI” (Birds)
Volume: 25 (2015/1)
Year: 2015
Pages: 20-23
Country: Lithuania

Contribution: photos in article about differences between Common Gull and Ring-billed Gull, based on my blog post Ring-billed Gulls vs Common Gulls: comparison from a Dutch perspective.


Publication: De Grauwe Gors (Avifauna Groningen)
Volume: 41-2013
Year: 2013
Page: 50
Country: The Netherlands

Title: Spaanse Kokmeeuwen in Noord-Nederland
(Spanish Black-headed Gulls in the North of Holland.)
Author: Klaas van Dijk

Contribution: 1 colour photograph


Publication: Aves
Volume: 49/4
Year: 2012
Page range: 205 -224
Country: Belgium

Title: Statut du Goéland brun Larus fuscus dans l’Entre-Sambre-et-Meuse : déplacements, origine, effectifs… Importance du site des barrages de l’Eau d’Heure
(Status of Lesser Black-backed Gull Larus fuscus in the Entre-Sambre-et-Meuse (Belgium): movements, provenance, numbers.)
Authors: Philippe Deflorenne & Fanny Carion Ellis

Contribution: 3 colour photos (pages 207, 220 and 223)

Sula 23(2)

Publication: SULA
Volume: 23(2)
Year: 2010
Page range: 69-86
Country: The Netherlands

Title: Rui en plaatstrouw van Zilvermeeuw Larus argentatus ZDGA op grond van kleurringaflezingen
(Moult and site-fidelity of Herring Gull ‘ZDGA’ based on colour-ring sightings)
Authors: Maarten van Kleinwee & Kees (C.J.) Camphuysen

Contribution: text and photographs.




Title: The Kittiwake
Publisher: Bloomsbury Publishing
Year: 2011
Country: United Kingdom
Author: John C. Coulson

Contribution: 2 colour photos


Title: National Geographic Bird-Watcher’s Bible
Publisher: National Geographic
Year: 2012
Country: USA

Contribution: one colour photo of a Mediterranean Gull (page 120).


Cover of the thesis. Copyright NIOZ.

Cover of the thesis. Copyright NIOZ.

Title: A historical ecology of two closely related gull species (Laridae): multiple adaptations to a man-made environment
Author: Kees Camphuysen
Year: 2013
Country: The Netherlands

Contribution: various photos

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